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The HR1 region constitutes the main helical stalk of S2, whereas the HR2 region is temporarily flexible in the prefusion state

Its biological functions extend beyond direct free-radical scavenging

J Basic Microbiol 50:494498 Mehdi K, Thierie J, Penninckx MJ (2001) gamma-Glutamyl transpeptidase in the yeast Saccharomyces cerevisiae and its role in the vacuolar transport and metabolism of glutathione