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Kla was enriched on cytoplasmic/mitochondrial proteins and clustered in central metabolic pathways (glycolysis/TCA, nucleotide metabolism), with lactylated hub enzymes (e.g., GAPDH, LDHA, ATP5F1A/B) anchoring a dense metabolic interaction network, positioning Kla as an organized regulator of carbon metabolism and a potential therapeutic vulnerability in ccRCC [280]

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